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A websites navigation is a critical part of it's usability and accessibility to spiders. A site with an effective and easy to use navigation makes it easy for visitors to find the pages that interest them, and keeps them on track, so they don't get lost. An effective navigation should make it obvious where a visitor is currently and how that location fits into the rest of the site structure.
In order to promote a search spider to crawl and index as much of your site as possible, your navigation should also be search engine friendly.